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  1. Wallwork, 1983 — Wallwork, J.A., 1983. Earthworm Biology. Edward Arnold, London, 58 pp.
  2. Wampler, 1982 — Wampler, J.E., 1982. The bioluminescence system of Microscolex phosphoreus and its similarities to those of other bioluminescent earthworms (oligochaeta). Comparative Biochemistry and Physiology Part A: Physiology, 71(4): 599-604.
  3. Wampler and Jamieson, 1980 — Wampler, J.E., Jamieson, B.G.M., 1980. Earthworm bioluminescence: comparative physiology and biochemistry. Comparative Biochemistry and Physiology, 66B: 43-50.
  4. Wampler and Jamieson, 1986 — Wampler, J.E., Jamieson, B.G.M., 1986. Cell bound bioluminescence from Pontodrilus bermudensis, and its similarities to other earthworm bioluminescence. Comparative Biochemistry and Physiology, 84A: 81-87.
  5. Wang et al., 2012 — Wang, J.H., Zhu, L.S., Liu, W., Wang, J., Xie, H., 2012. Biochemical responses of earthworm (Eisenia foetida) to the pesticides chlorpyrifos and fenvalerate. Toxicol. Mech. Methods, 22(3): 236-241.
  6. Wang H.J. and Qiu, 2005 — Wang, H.J., Qiu, J.P., 2005. A Preliminary Report on Pheretimoid Earthworms from the Xingdou Mountain Nature Reserve-With Description of Two New Species and a New Subspecies of the Genera Amynthas and Metaphire. Journal of Shanghai Jiaotong University (Agricultural Science), 23(1): 23-30. [In Chinese with English summary].
  7. Wang H.Z. and Cui, 2007 — Wang, H.Z., Cui, Y.D., 2007. On the studies of Microdrile Oligochaeta and Aeolosomatidae (Annelida) in China: brief history and species checklist. Acta Hydrobiologica Sinica, 31: 87-98.
  8. Wang K.Y. et al., 2013 — Wang, K.Y., Tull, L., Cooper, E., Wang, N., Liu, D., 2013. Recombinant Protein Production of Earthworm Lumbrokinase for Potential Antithrombotic Application. Evidence-Based Complementary and Alternative Medicine, Article ID 783971, 8 pp., doi.org/10.1155/2013/783971.
  9. Wang Y. et al., 2012 — Wang, Y., Wu, S., Chen, L., Wu, C., Yu, R., Wang, Q., Zhao, X., 2012. Toxicity assessment of 45 pesticides to the epigeic earthworm Eisenia fetida. Chemosphere, 88(4): 484-491.
  10. Wang Y.H. and Shih, 2010 — Wang Y.H., Shih, H.T., 2010. Earthworm fauna of Eastern Taiwan, with descriptions of two new species (Oligochaeta: Megascolecidae). Zootaxa, 2341: 52–68.
  11. Wang Y.H. and Shih, 2017 — Wang Y.H., Shih, H.T., 2017. Four New Species of Earthworms (Oligochaeta: Megascolecidae: Amynthas) from Taiwan Based on Morphological and Molecular Evidence. Zoological Studies, 56: 1-12, doi:10.6620/ZS.2017.56-18.
  12. Warnick, 1978 — Warnick, J.W., 1978. Use of the earthworm Eiseniella tetraedra as a bioassay test organism. Deep Blue at the University of Michigan.
  13. Watanabe and Olson, 1965 — Watanabe, F.S., Olson, S.R., 1965. Test of an ascorbic acid method for determining phosphorus in water and NaHCO 3 extracts from soils. Soil Science Society of America Proceedings, 29: 677–678.
  14. Watanabe and Tsukamoto, 1976 — Watanabe, H., Tsukamoto, J., 1976. Seasonal change in size, class and stage structure of lumbricid Eisenia foetida population in a field compost and its practical application as the decomposer of organic waste matter. Rev. Ecol. Biol. Sol., 13: 141-146.
  15. Watanabe et al., 1966 — Watanabe, H., Saichuae, P., Shidei, T., 1966. On the biomass of soil animals found in various types of forests in Thailand [On-line]. Available http://hdl.handle.net/2433/55202.
  16. Way and Scopes, 1968 — Way, M.J., Scopes, N.E.A., 1968. Studies on the persistence and effects on soil fauna of some soil-applied systemic insecticides. Ann. Appl. Biol., 62: 199–214.
  17. Webster, 1895 — Webster, F.M., 1895. The Earthworm as a tree-planter. J. N. York Ent. Soc., III: 139.
  18. Weltje, 1998 — Weltje, L., 1998. Mixture toxicity and tissue interactions of Cd, Cu, Pb and Zn in earthworms (Oligochaeta) in laboratory and field soils, a critical evaluation of data. Chemosphere, 36: 2643–2660.
  19. Werner and Dindal, 1989 — Werner, M.R., Dindal, D.L., 1989. Earthworm community dynamics in conventional and low-input agroecosystems. Rev. Écol. Biol. Sol., 26(4): 427–437.
  20. Wessely, 1905 — Wessely, K., 1905. Die Lumbriciden Oberösterreichs. Jahresber. Ver. Nat. Linz, XXXIV, 19 p.
  21. Wetzel, 2003 — Wetzel, M.J., 2003. The terrestrial Oligochaeta of Illinois - Annotated checklist of species. Illinois Natural History Survey, University of Illinois.
  22. Wetzel, 2004 — Wetzel, M.J., 2004. The terrestrial Oligochaeta of Illinois - Annotated checklist of species. Illinois Natural History Survey, University of Illinois.
  23. Wetzel, 2005 — Wetzel, M.J., 2005. Checklist of the aquatic and terrestrial Oligochaeta occuring in North Carolina, South Carolina, and/or Tennessee, with notations for species now known to occur in the Great Smocky Mountains National Park. Illinois Natural History Survey, University of Illinois.
  24. Wetzel, 2007 — Wetzel, M.J., 2007. The terrestrial Oligochaeta of Illinois - Annotated checklist of species. Illinois Natural History Survey, University of Illinois.
  25. Wetzel and Taylor, 2001 — Wetzel, M.J., Taylor, S.J., 2001. First records of freshwater oligochaetes (Annelida, Clitellata) from caves in Illinois and Missouri, USA. Journal of Cave and Karst Studies, 63(3): 99-104.
  26. Whalen and Parmelee, 1999 — Whalen, J.K., Parmelee, R.W., 1999. Growth of Aporrectodea tuberculata (Eisen) and Lumbricus terrestris L. under laboratory and field conditions. Pedobiologia, 43: 1–10.
  27. Whalen and Parmelee, 2000 — Whalen, J.K., Parmelee, R.W., 2000. Earthworm secondary production and N flux in agroecosystems: a comparison of two approaches. Oecologia, 123.
  28. Whalen et al., 1998 — Whalen, J.K., Parmelee, R.W., Edwards, C.A., 1998 Population dynamics of earthworm communities in corn agroecosystems receiving organic or inorganic fertilizer amendments. Biol. Fertil. Soils, 27: 400–407.
  29. Whalen et al., 1999 — Whalen, J.K., Parmelee, R.W., McCartney, D.A., VanArsdale, J.L., 1999. Movement of N from decomposing earthworm tissue to soil, microbial and plant N pools. Soil Biol. Biochem., 31: 487–492.
  30. Whalen et al., 2000 — Whalen, J.K., Parmelee, R.W., Subler, S., 2000. Quantification of nitrogen excretion rates for three lumbricid earthworms using 15N. Biol. Fertil. Soils, 32: 347–352.
  31. Wijnhoven, 2006 — Wijnhoven, S., Thonon, I., van der Velde, G., Leuven, R., Zorn, M., Eijsackers, H., et al., 2006. The impact of bioturbation by small mammals on heavy metal redistribution in an embanked floodplain of the river Rhine. Water Air Soil Pollut., 177: 183–210.
  32. Wilcke, 1952 — Wilcke, D.E., 1952. Zur Kenntnis der Lumbricidenfaunen Deutschlands. Zool. Anz., 151: 104–106.
  33. Willem and Minue, 1900 — Willem, V., Minue, A., 1900. La signification des cellules jaunes de l'intestin du Lombric. C. R. Ass. France, XXVIII: 271.
  34. Wills and Abbott, 2003 — Wills, A., Abbott, I., 2003. Landscape-scale species richness of earthworms in the Porongurup Range, Western Australia: influence of aspect, soil fertility and vegetation type. Biology and Fertility of Soils, 39: 94–102.
  35. Wilson, 1961 — Wilson, D.M., 1961. The connections between the lateral giant fibers of earthworms. Comparative Biochemistry and Physiology, 3(4): 274–284.
  36. Wilson W.J. et al., 2014 — Wilson, W.J., Ferrara, N.C., Blaker, A.L., Giddings, C.E., 2014. Escape and avoidance learning in the earthworm Eisenia hortensis. Peer J., 2: e250.
  37. Winsome et al., 2006 — Winsome, T., Hendrix, P.F., Epstein, L., Horwath, W.R., 2006. Competitive interactions between native and exotic earthworm species as influenced by habitat quality in a California grassland. Appl. Soil Ecol., 32: 38-53.
  38. Winterbourn, 1978 — Winterbourn, M.J., 1978. The macroinvertebrate fauna of a New Zealand forest stream. New Zealand Journal of Zoology, 5: 157-169.
  39. Wironen and Moore, 2006 — Wironen, M., Moore., T.R., 2006. Exotic earthworm invasion increases soil carbon and nitrogen in an old-growth forest in southern Quebec. Can. J. For. Res., 36: 845–854.
  40. Wolff et al., 2006 — Wolff, W.J., Duiven, A.G., Duiven, P., Esselink, P., Gueye, A., Meijboom, A., Moerland, G., Zegers, J., 1993. Biomass of macrobenthic tidal flat fauna of the Banc d'Arguin, Mauritania. Hydrobiologia, 258: 151-163, doi:10.1007/BF00006193.
  41. Wolters, 1991 — Wolters, V., 1991. Soil invertebrates - effects on nutrient turnover and soil structure - a review. Zeitschrift für Pflanzenernährung und Bodenkunde, 154: 389-402.
  42. Wolters and Stickan, 1991 — Wolters, V., Stickan, W., 1991. Resource allocation of beech seedlings (Fagus sylvatica L.) relationship to earthworm activity and soil conditions. Oecologia, 88(1): 125-131.
  43. Wolters and Stickan, 1991b — Wolters, V., Stickan, W., 1991. Effects of an endogeic earthworm species on transpiration and C-allocation of beech (Fagus sylvatica L.) seedlings. Ber. Forschungszentrums Waldökosysteme (B), 22: 383–386.
  44. Wolters and Joergensen, 1992 — Wolters, V., Joergensen, R.G., 1992. Microbial carbon turnover in beech forest soils worked by Aporrectodea caliginosa (Savigny) (Oligochaeta: Lumbricidae). Soil Biol. Biochem., 24: 171-177.
  45. Wolters and Schaefer, 1993 — Wolters, V., Schaefer, M., 1993. Effects of burrowing by the earthworm Aporrectodea caliginosa (Sabigny) on beech litter decomposition in an agricultural and in a forest soil. Geoderma, 56: 627-632.
  46. Wolters and Ekschmitt, 1995 — Wolters,V., Ekschmitt, K., 1995. Earthworm effects on the use of C sources by microorganisms: Non-linear response to temperature alteration. Biology and Fertility of soils, 19: 109-114.
  47. Wood and James, 1993 — Wood, H.B., James, S.W., 1993. Native and introduced earthworms from selected chaparral, woodland, and riparian zones in southern California. Gen.Tech. Rep. PSW-GTR-142. Albany, CA: Pacific Southwest Research Station, Forest Service, USDA, 20 pp.
  48. Wood et al., 1997 — Wood, H.B., Olivier, K.L., James, S.W., 1997. Relict Megascolecidae and exclusion of Lumbricidae from basalt-derived soils in Southern California. Soil Biology and Biochemistry, 29: 241-244.
  49. Wroot, 1985 — Wroot, A.J., 1985. A Quantitative Method for Estimating the Amount of Earthworm (Lumbricus terrestris) in Animal Diets. Oikos, 44(2): 239-242.
  50. Wu and Sun, 1997 — Wu, J., Sun, X., 1997. On one species of Amynthas (Oligochaeta: Megascolecidae) from China. Sichuan Journal of Zoology, 1997-01.
  51. Wyssotzky, 1898 — Wyssotzky, G., 1898. Der Boden, der Grund und die Hydrostatica des Welikoanadoljschen Reviers]. Truduiete. [Arbeiten der vom Walddepartement unter Prof. Dokutschajeff's Leitung ausgerüsteten Expedition; Natur- und Pflanzenkultur im Weliko-anadoljschen Revier]. St. Petersburg. 94 p. [In Russian].