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Contents

Acanthodrilidae (Claus, 1880)

Acanthodrilinae (Claus, 1880)

Acanthodrilus (Perrier, 1872)

Bahlia

Barogaster (Gates, 1940)

Benhamiinae (Michaelsen, 1897)

Agastrodrilus

Balanta (Michaelsen, 1898)

Benhamia

Benhamiona (Csuzdi and Zicsi, 1994)

Borgesia (James, 1991)

Calebiella (Gates, 1945)

Chaetocotoides (Julka, 1988)

Chilota (Michaelsen, 1899)

Cubadrilus (Rodriguez Aragones and Carlos, 2002)

Dashiella (Julka, 1988)

Decachaetus (Lee, 1959)

Deinodrilus (Beddard, 1888)

Deltania (Eisen, 1893)

Dichogaster (Beddard, 1888)

Dinodriloides (Benham, 1904)

Dinodrilus

Diplocardia (Garman, 1888)

Diplotrema (Spencer, 1900)

Dudichiodrilus (Csuzdi, 1995)

Eodriloides (Zicsi, 1998)

Eodrilus (Michaelsen, 1907)

Erythraeodrilus

Eudichogaster (Michaelsen, 1902)

Eudinodriloides (Lee, 1959)

Eudiplotrema

Eutrigaster

Eutyphoeus (Michaelsen)

Exxus

Geodrilus (Ude, 1893)

Guineoscolex (Csuzdi and Zicsi, 1994)

Herbettodrilus (Julka, Blanchart and Chapuis-Lardy, 2004)

Hoplochaetella (Michaelsen, 1900)

Hoplochaetina (Michaelsen, 1926)

Karmiella (Julka, 1983)

Kaxdrilus (Fragoso and Rojas-Fernandez, 1994)

Kayarmacia (Jamieson, 1997)

Konkadrilus (Julka, 1988)

Kotegeharia (Julka, 1988)

Larsonidrilus (James, 1994)

Lavellodrilus (Fragoso, 1988)

Leptodrilus (Benham, 1909)

Leucodrilus (Lee, 1952)

Loksaia (Csuzdi, 1996)

Maheina (Michaelsen, 1899)

Mallehulla (Julka, 1982)

Mandane (Kinberg, 1867)

Maoridrilus (Michaelsen, 1899)

Mayadrilus (Fragoso and Rojas-Fernandez, 1994)

Microdrilus (Beddard, 1893)

Microscolex (Rosa, 1887)

Millsonia

Monogaster (Michaelsen, 1915)

Monothecodrilus (Csuzdi and Zicsi, 1994)

Neochaeta (Lee, 1959)

Neodiplotrema (Dyne, 1997)

Neodrilus (Beddard, 1887)

Neogaster (Cernosvitov, 1934)

Neotrigaster (James, 1991)

Notiodrilus (Michaelsen, 1899)

Octochaetoides

Octochaetona (Gates, 1962)

Octochaetus (Beddard, 1893)

Octonochaeta (Julka, 1988)

Omodeona (Sims, 1967)

Parachilota (Pickford, 1937)

Parryodrilus (Julka, Giri, Panigrahi and Senapati, 1997)

Pellogaster (Gates)

Perieodrilus (Michaelsen, 1910)

Photodrilus (Giard, 1887)

Pickfordia

Plagiochaeta (Benham, 1892)

Plutellus (Perrier, 1873)

Priodochaeta

Priodoscolex

Protozapotecia (James, 1994)

Ramiella (Stephenson, 1921)

Reginaldia (Csuzdi, 2006)

Peritogaster (Csuzdi, 2006)

Reginaldia (Csuzdi, 2006)

Rhododrilus (Beddard, 1889)

Rillogaster (Gates, 1939)

Senapatiella (Julka, Blanchart and Chapuis-Lardy, 2004)

Shimodrilus (Julka, Blanchart and Chapuis-Lardy, 2004)

Sylvodrilus (Lee, 1959)

Torresiella (Dyne, 1997)

Trigaster (Benham, 1886)

Typhoeus

Udeina (Michaelsen, 1910)

Wahoscolex (Julka, 1988)

Wegeneriella

Wegeneriona

Yagansia (Michaelsen, 1899)

Balanteodrilus (Pickford, 1938)

Ramiellona

Zapatadrilus (James, 1991)

Zapotecia (Eisen, 1900)

Ailoscolecidae (Bouché, 1969)

Ailoscolex (Bouché, 1969)

Almidae (Duboscq, 1902)

Alma (Grube, 1855)

Areco (Righi, Ayres and Bittencourt, 1978)

Drilocrius (Michaelsen, 1917)

Glyphidrilus (Horst, 1889)

Progizzardus (Nair, 2010)

Biwadrilidae (Vejdovsky, 1884)

Biwadrilus (Brinkhurst and Jamieson, 1971)

Criodrilidae (Vejdovsky, 1884)

Criodrilus (Hoffmeister, 1845)

Eudrilidae (Claus, 1880)

Eudrilinae (Claus, 1880)

Beddardiella (Michaelsen, 1910)

Bettonia (Beddard, 1903)

Bettoniella (Gates, 1941)

Borgertia (Michaelsen, 1905)

Buettneriodrilus (Michaelsen, 1897)

Callidrilus (Michaelsen, 1890)

Chuniodrilus (Michaelsen, 1913)

Eminoscolex (Michaelsen, 1896)

Ephyriodrilus (Sims, 1971)

Eudriloides (Michaelsen, 1890)

Eudrilus (Perrier, 1871)

Eupolytoreutus (Michaelsen, 1910)

Euscolex (Michaelsen, 1902)

Eutoreutus (Michaelsen, 1915)

Gardullaria (Michaelsen, 1903)

Heliodrilus (Beddard, 1890)

Hippopera (Taylor, 1949)

Hyperiodrilus (Beddard, 1890)

Iridodrilus (Beddard, 1897)

Kaffania (Michaelsen, 1903)

Keffia (Clausen, 1965)

Lavellea (Csuzdi and Tondoh, 2007)

Legonea (Clausen, 1965)

Libyodrilus (Beddard, 1891)

Malodrilus (Michaelsen, 1903)

Metadrilus (Michaelsen, 1891)

Metascolex (Michaelsen, 1903)

Nemertodrilus (Michaelsen, 1890)

Neumanniella (Michaelsen, 1903)

Neumanniona (Michaelsen, 1935)

Nsukkadrilus (Segun, 1977)

Paraeudrilinae (Beddard, 1894)

Legonodrilus (Sims, 1969)

Parapolytoreutus (Segun, 1980)

Parascolex (Michaelsen, 1900)

Pareudrilus (Beddard, 1894)

Platydrilus (Michaelsen, 1891)

Polytoreutus (Michaelsen, 1890)

Reithrodrilus (Michaelsen, 1891)

Rosadrilus (Cognetti de Martiis, 1909)

Schoutedenina (Michaelsen, 1936)

Schubotziella (Michaelsen, 1915)

Scolecillus (Omodeo, 1958)

Segunia (Sims, 1985)

Segunodrilus (Owa, 1995)

Stuhlmannia (Michaelsen, 1890)

Takunodrilus (Owa, 1995)

Teleudrilus (Rosa, 1888)

Teleutoreutus (Michaelsen, 1903)

Tubiscolex (Michaelsen, 1935)

Vomia (Segun, 1976)

Alvania

Notykus (Michaelsen, 1891)

Unyoria (Michaelsen, 1896)

Glossoscolecidae (Michaelsen, 1900)

Diaguita (Cordero, 1942)

Enantiodrilus (Cognetti de Martiis, 1902)

Eurydame (Kinberg, 1867)

Fimoscolex (Michaelsen, 1900)

Glossodrilus (Cognetti de Martiis, 1905)

Glossoscolex (Leuckart, 1835)

Holoscolex (Cognetti de Martiis, 1904)

Righiodrilus (Zicsi, 1995)

Titanus

Urochaeta

Haplotaxidae (Michaelsen, 1900)

Delaya (Brinkhurst, 1988)

Haplotaxis (Hoffmeister, 1843)

Hologynus (Brinkhurst, 1988)

Metataxis (Righi, 1985)

Nemodrilus (Claparède, 1862)

Omodeodrilus (Kammerer, 2006)

Pelodrilus (Beddard, 1891)

Hormogastridae (Michaelsen, 1900)

Hemigastrodrilus (Bouché, 1970)

Hormogaster (Rosa, 1887)

Vignysa (Bouché, 1970)

Xana (Diaz Cosin, Briones and Trigo, 1989)

Komarekionidae (Gates, 1974)

Komarekiona (Gates, 1974)

Kynotidae (Brinkhurst and Jamieson, 1971)

Kynotus (Michaelsen, 1891)

Lumbricidae (Rafinesque'Schmaltz, 1815)

Allolobophora (Eisen, 1873)

Allolobophoridella (Mrsic, 1990)

Aporrectodea (Örley, 1885)

Avelona (Qiu and Bouché, 1998)

Bimastos (Moore, 1893)

Cataladrilus (Qiu and Bouché, 1998)

Cernosvitovia (Omodeo, 1956)

Dendrobaena (Eisen, 1873)

Dendrodriloides (Kvavadze, 2000)

Dendrodrilus (Omodeo, 1956)

Diporodrilus (Bouché, 1970)

Eisenia (Malm, 1877)

Eiseniella (Michaelsen, 1900)

Eisenoides (Gates, 1969)

Eophila (Rosa, 1893)

Ethnodrilus (Bouché, 1972)

Eumenescolex (Qiu and Bouché, 1998)

Fitzingeria (Zicsi, 1978)

Healyella (Omodeo and Rota, 1989)

Helodrilus (Hoffmeister, 1845)

Heraclescolex (Qiu and Bouché, 1998)

Hydrilus (Qiu and Bouché, 1998)

Iberoscolex (Qiu and Bouché, 1998)

Italobalkaniona (Mrsic and Sapkarev, 1988)

Karpatodinariona (Mrsic and Sapkarev, 1988)

Koinodrilus (Qiu and Bouché, 1998)

Kritodrilus (Bouché, 1972)

Lumbricus (Linnaeus, 1758)

Microeophila (Omodeo, 1956)

Murchieona (Gates, 1978)

Norealidys

Octodriloides (Zicsi, 1986)

Octodrilus (Omodeo, 1956)

Octolasion (Örley, 1885)

Orodrilus (Bouché, 1972)

Perelia (Easton, 1983)

Pietromodeona (Qiu and Bouché, 1998)

Postandrilus (Qiu and Bouché, 1998)

Proctodrilus (Zicsi, 1985)

Prosellodrilus (Bouché, 1972)

Riphaeodrilus (Csuzdi and Pavlicek, 2005)

Satchellius (Gates, 1975)

Scherotheca (Bouché, 1972)

Spermophorodrilus (Bouché, 1975)

Zophoscolex (Qiu and Bouché, 1998)

Lutodrilidae (McMahan, 1976)

Lutodrilus (McMahan, 1976)

Megascolecidae (Rosa, 1891)

Aceeca (Blakemore, 2000)

Amphimiximus (Blakemore, 2000)

Amynthas (Kinberg, 1867)

Anisochaeta (Beddard, 1890)

Anisogaster (Blakemore, 2000)

Aporodrilus (Blakemore, 2000)

Archipheretima (Michaelsen, 1928)

Arctiostrotus (McKey-Fender)

Argilophilus (Eisen, 1893)

Austrohoplochaetella (Jamieson, 1971)

Begemius (Easton, 1982)

Caecadrilus (Blakemore, 2000)

Celeriella (Gates, 1958)

Chetcodrilus (Fender and McKey-Fender, 1990)

Comarodrilus (Stephenson, 1915)

Cryptochaeta

Cryptodrilus (Fletcher, 1886)

Dendropheretima (James, 2004)

Deodrilus (Beddard, 1890)

Didymogaster (Fletcher, 1886)

Digaster (Perrier, 1872)

Diporochaeta (Beddard, 1890)

Drilochaera

Driloleirus (Fender and McKey-Fender, 1990)

Duplodicodrilus (Blakemore, 2008)

Eastoniella (Jamieson, 1977)

Fletcherodrilus (Michaelsen, 1891)

Gastrodrilus (Blakemore, 2000)

Gemascolex (Edmonds and Jamieson, 1973)

Graliophilus (Jamieson, 1971)

Haereodrilus (Dyne in Jamieson, 2000)

Heterochaetella (Yamaguchi, 1953)

Heteroporodrilus (Jamieson, 1970)

Hiatidrilus (Jamieson, 1994)

Hickmaniella (Jamieson, 1974)

Howascolex

Hypolimnus (Blakemore, 2000)

Isarogoscolex (James, 2004)

Kanchuria (Julka, 1988)

Kincaidodrilus (McKey-Fender, 1982)

Lampito (Kinberg, 1867)

Lennogaster (Gates, 1939)

Macnabodrilus (Fender and McKey-Fender, 1990)

Mahbenus (Bourne, 1894)

Megascolex (Templeton, 1844)

Megascolides (McCoy, 1878)

Metapheretima (Michaelsen, 1928)

Metaphire (Sims and Easton, 1972)

Nellogaster (Gates, 1938)

Nelloscolex (Gates, 1939)

Nephrallaxis (Fender and McKey-Fender, 1990)

Nexogaster (Blakemore and Kingston, 1997)

Notoscolex (Fletcher, 1886)

Occicryptodrilus

Paraplutellus (Jamieson, 1972)

Perichaeta (Schmarda, 1861)

Pericryptodrilus (Jamieson, 1977)

Perionychella (Michaelsen, 1907)

Perionyx (Perrier, 1872)

Perissogaster (Fletcher, 1887)

Pheretima (Kinberg, 1867)

Pithemera (Sims and Easton, 1972)

Planapheretima (Michaelsen, 1934)

Pleionogaster (Michaelsen, 1892)

Pleurochaeta (Beddard, 1883)

Plutelloides (Jamieson, 2000)

Polypheretima (Michaelsen, 1934)

Pontodrilus (Perrier, 1874)

Propheretima (Jamieson, 1995)

Provescus (Blakemore, 2000)

Pseudocryptodrilus (Jamieson, 1972)

Pseudonotoscolex (Jamieson, 1971)

Pseudoperichaeta (Jamieson, 1970)

Reflechtodrilus (Blakemore, 2005)

Retrovescus (Blakemore, 1998)

Scolecoidea (Blakemore, 2000)

Scolioscolides (Gates, 1937)

Sebastianus (Blakemore, 1997)

Simsia (Jamieson, 1972)

Spenceriella (Michaelsen, 1907)

Tassiedrilus (Blakemore, 2000)

Terriswalkerius (Jamieson, 1994)

Tokea (Benham, 1904)

Tonoscolex (Gates, 1933)

Toutellus (Fender and McKey-Fender, 1990)

Travoscolides (Gates, 1940)

Trinephrus (Beddard, 1895)

Troyia (Jamieson, 1977)

Vesiculodrilus (Jamieson, 1973)

Woodwardiella (Stephenson, 1925)

Zacharius (Blakemore, 1997)

Microchaetidae (Michaelsen, 1900)

Brachydrilus (Benham, 1888)

Microchaetus (Rapp, 1849)

Proandricus (Plisko, 1992)

Moniligastridae (Claus, 1880)

Desmogaster (Rosa, 1890)

Drawida (Michaelsen, 1900)

Eupolygaster (Michaelsen, 1900)

Hastirogaster (Gates, 1930)

Moniligaster (Perrier, 1872)

Ocnerodrilidae (Beddard, 1891)

Malabarinae (Gates, 1966)

Malabaria (Stephenson, 1924)

Nannodrilus (Beddard, 1894)

Nematogenia (Eisen, 1900)

Ocnerodrilinae (Beddard, 1891)

Aphanascus (Stephenson, 1924)

Bauba (Righi, 1980)

Belladrilus (Righi, 1984)

Chacdrilus (Fragoso and Rojas, 2009)

Curgiona (Gates, 1941)

Dariodrilus (Righi, Ayres and Bittencourt, 1978)

Deccania (Gates, 1949)

Diaphorodrilus (Cognetti de Martiis, 1910)

Dorgiodrilus (Gates, 1962)

Enicmodrilus (Eisen, 1900)

Eukerria (Michaelsen, 1935)

Exisdrilus (Righi, Ayres and Bittencourt, 1978)

Filodrilus (Chen, 1938)

Gatesia (Jamieson, 1962)

Gordiodrilus (Beddard, 1892)

Haplodrilus (Eisen, 1900)

Ilyogenia (Beddard, 1893)

Kerria

Kerriona (Michaelsen, 1926)

Liodrilus (Eisen, 1900)

Lourdesia (Righi, 1994)

Ocnerodrilus (Eisen, 1878)

Paulistus (Michaelsen, 1925)

Phoenicodrilus (Eisen, 1895)

Pygmaeodrilus (Michaelsen, 1890)

Quechuona (Gates, 1941)

Tazelaaria (Gates, 1962)

Thatonia (Gates, 1942)

Xibaro (Righi, 1981)

Pontoscolecidae (James, 2012)

Aicodrilus (Righi, 1971)

Alexidrilus (Righi, 1971)

Andiodrilus (Michaelsen, 1900)

Andiorrhinus (Cognetti de Martiis, 1908)

Andioscolex (Michaelsen, 1927)

Anteoides (Cognetti de Martiis, 1902)

Anteus (Perrier, 1872)

Aptodrilus (Cognetti de Martiis, 1904)

Atatina (Righi, 1971)

Aymara (Michaelsen, 1935)

Botarodrilus (Zicsi, 1990)

Bribri (Righi, 1984)

Chibui (Righi and Guerra, 1985)

Cirodrilus (Righi, 1975)

Diachaeta (Benham, 1886)

Estherella (Gates, 1970)

Eudevoscolex (Cordero, 1944)

Goiascolex (Righi, 1971)

Hexachyloscolex (Zicsi and Csuzdi, 1999)

Inkadrilus (Michaelsen, 1917)

Langioscolex (Zicsi, 1990)

Martiodrilus (Michaelsen, 1937)

Meroscolex (Cernosvitov, 1934)

Nouraguesia (Csuzdi and Pavlicek, 2011)

Onoreodrilus (Zicsi, 1988)

Onychochaeta (Beddard, 1891)

Opisthodrilus (Rosa, 1895)

Periscolex (Cognetti de Martiis, 1905)

Perolofius (Righi and Nemeth, 1983)

Pontoscolex (Schmarda, 1861)

Pseudochibui (Drachenberg, 1991)

Quimbaya (Michaelsen, 1935)

Randdrilus (Moreno and Perez-Santos, 1997)

Rhinodrilus (Perrier, 1872)

Tairona (Righi, 1984)

Tamayodrilus (Zicsi, 1995)

Thamnodriloides (Gates, 1968)

Thamnodrilus (Beddard, 1887)

Tuiba (Righi, Ayres and Bittencourt, 1976)

Tupinaki (Righi, 1995)

Urobenus (Benham, 1886)

Zongodrilus (Righi, 1996)

Sparganophilidae (Michaelsen, 1918)

Sparganophilus (Benham, 1892)

Syngenodrilidae (Smith and Green, 1919)

Syngenodrilus (Smith and Green, 1919)

Tiguassuidae (Brinkhurst, 1988)

Tiguassu (Righi, Ayres and Bittencourt, 1978)

Tritogeniidae (Plisko, 2013)

Michalakus (Plisko, 1996)

Tritogenia (Kinberg, 1867)

Tumakidae (Righi, 1995)

Tumak (Righi, 1996)

Digitibranchus (Levinsen, 1889)

Geoscolex (Leuckart, 1841)

Irewiae

Siphonogaster

Tykonus (Michaelsen, 1892)